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Plant bio
Water and solute movement
Question | Answer |
---|---|
How does water move? | Transpiration Water vapour pressure (deficit)/relative humidity Water potential Solute potential Turgor pressure (potential) Matrix potential Resistance Surface tension |
Transpiration | Water movement through the plant Water (liquid) column is continuous from soil water to leaf tissues These continuities persist in growth & development Most water is lost as vapour/must be replaced Phase change at tissue/air space boundary |
Transpiration and plant water potential | - plant in wet soil, stomata closed: water potential zero (high), atmosphere low - stomata opens: water potential inside falls as water vapour leaves - difference is proportional to resistance (sum of all) |
Liquid flow resistances | Lack of resistance means flow is an instantaneous response to differences in water potential - resistance across xylem is greater than along leads to differences in water potential from one side to another |
Stem resistances | Stem resistances are low Highest resistance in petioles – this isolates leaves from each other Resistances give rise to pressure differences |
Water in xylem | - under tension - evidence of negative potentials in xylem |
Absorption lag | Consider a plant transpiring, if you stop transpiration (by immersing leaves) what happens? - Plant carries on absorbing water, with a lag; but at diminishing rate, biphasic nature indicates different routes with different resistances |
Water potential Ψ | the forces acting on water In plants, water moves from areas of +ve potential to -ve potential is less (more –ve): under hydrostatic tension, in osmotic solution or on adsorption to surface Cell water potential = pressure potential + osmotic potential |
Water in cells and tissues | Consider cell, with overall water potential = -5.0 bar, osmotic potential Ψsap = -5 bar, pressure potential Ψp = 0 bar and cell wall with water potential Ψ= -1 bar water flows into cell (and hence vacuole) from cell wall with pressure of 4 bar |
Causing Ψsap to increase (become less –ve) Thus, difference Ψcell vs Ψwall reduces, and Ψp (of cell) increases (becomes +ve) | |
Water transport in xylem | Water movement through the xylem requires ten times less pressure than movement through living cells - simple low resistance pathway |
Cohesion tension theory | Tension developed at the top of the tree pulls water through the xylem: requires cohesive properties of water to sustain large tensions in the xylem water columns |
Water is brought to the leaves via the xylem of the leaf vascular bundle, branched into a very fine network of veins Most cells in a typical leaf are within 0.5 mm of a minor vein, water is drawn into the cells of the leaf and along the cell walls | |
Evaporation | Water evaporation from the leaf air spaces generates a negative pressure in the xylem |
Translocation in phloem | The phloem is the tissue that translocates the products of photosynthesis from mature leaves to areas of growth and storage, including the roots Sugar is translocated in phloem sieve elements |
Pathways of translocation | Mature sieve elements are living cells highly specialised for translocation |
Materials translocated in phloem | Sucrose, at rate 104 x that in other symplasts Amino acids Hormones Inorganic ions RNA |
Phloem loading | - sugar from phloem to sink cells - from chloroplasts to sieve elements - occurs from mesophyll cells via the apoplast or the symplast (different companion cells are used) |
Pressure flow model | - phloem translocation as a flow of solution driven by an osmotically generated pressure gradient between source and sink |