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Touch

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Question
Answer
Kinesthesis   perception of the position & movement of our limbs in space  
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Proprioception   perception mediated by kinesthetic & vestibular receptors  
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Somatosensation   collective term for sensory signals frm the body  
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Epidermis   outer of 2 major layers of skin  
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Dermis   inner of 2 major layers of skin, consisting of nutritive & connective tissues, w/in which lie the mechanoreceptors  
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Mechanoreceptors   sensory receptors that are responsive to mechanical stimulation (pressure, vibration, & movement)  
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Meissner corpuscle   specialized nerve ending associated w/fast-adapting (FA 1) fibers that have small receptive fields  
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Merkel cell neurite complex   specialized nerve ending assoc. w/slow-adapting (SA 1) fibers that have small receptive fields  
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Pacinian corpuscle   specialized nerve endind assoc. w/fast-adapting (FA 2) fibers that have large receptive fields  
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Ruffini ending   specialized nerve ending assoc. w/slow-adapting (SA 2) fibers that have large receptive fields  
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Kinesthetic   referring to perception involving sensory mechanoreceptors in muscles, tendons, & joints  
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Muscle spindle   sensory receptor located in a muscle that senses its tension  
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Thermoreceptors   sensory receptors that signal info. abt changes in skin temperature  
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Warmth fiber   sensory nerve fiber that fires when skin temp. increases  
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Cold fiber   sensory nerve fiber that fires when skin temp. decreases  
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Nocioreceptors   sensory receptors that transmit info. abt noxious (painful) stimulation that causes damage or potential damage to the skin  
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A-delta fiber   intermediate-sized, myelinated sensory nerve fiber that transmits pain and temp. signals  
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C-fiber   narrow-diameter, unmyelinated sensory nerve fiber that transmits pain and temp. signals  
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Spinothalamic pathway   route from the spinal cord to the brain that carries most of the info. abt skin temp. and pain  
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Dorsal column-medial lemniscal (DCML) pathway   route frm the spinal cord to the brain that carries signals frm skin, muscles, tendons, & joints  
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Somatosensory area 1(S1)   primary receiving area for touch in the cortex; located in the parietal lobe  
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Somatosensory area 2 (S2)   secondary receiving area for touch in the cortex; located in the upper bank of lateral sulcus & other cortical areas  
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Somatotopic   spatially mapped in the somatosensory cortex in correspondence to spatial events in the skin  
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Homunculus   maplike representation of regions of the body in the brain  
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Phantom limb   sensation perceived frm a physically amputated limb of the body  
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Neural plasticity   ability of neural circuits to undergo changes in function or organization as a result of previous activity  
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Substantia gelatinosa   jellylike region of interconnecting neurons in the dorsal horn of the spinal cord  
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Dorsal horn   region @ the rear of the spinal cord that received input frm receptors in the skin  
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Gate control theory   description of the system that transmits pain that incorporates modulating signals frm the brain  
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Anterior cingulate cortex (ACC)   region of the brain assoc. with the perceived unpleasantness of a pain sensation  
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Prefrontal cortex (PFC)   region of the brain concerned w/cognitive and executive control  
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Analgesia   decreasing pain sensation during conscious experience  
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Endogenous opiates   chemicals released by the body that block the release or uptake of neurotransmitters necessary to transmit pain sensation to the brain  
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Hyperalgesia   increased on heightened response to a normally painful stimulus  
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Two-point threshold   min. distance @ which 2 stimuli (eg. 2 simultaneous touches) are just perceptible as separate  
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Haptic perception   knowledge of the world that is derived frm sensory receptors in the skin, muscles, tendons, & joints, usually involving active exploration  
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Exploratory procedure   stereotyped hand movement patterns used to contact objects in order to perceive their properties; each exploratory procedure is best for determining 1(or more) object properties  
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Tactile agnosia   inability to identify objects by touch  
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Frame of reference   coordinate system used to define locations in space  
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Egocenter   center of reference frame used to represent locations relative to the body  
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Body image   the impression of our bodies in space  
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Endogenous spatial attention; form of top-down control (knowledge-driven) of spatial attn   spatial attn. in which attn. is voluntarily directed toward the site where the observer anticipates a stimulus will occur  
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Exogenous spatial attention; form of bottom-up (stimulus-driven) spatial attn. in which attn. is reflexively directed toward the site @ which a stimulus has abruptly appeared    
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Tadoma   method by which those who are both deaf & blind can perceive speech in real time using their hands  
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Virtual haptic environment   synthetic world that may be experienced haptically by operation of an electromechanical device that delivers forces to the hand of the user  
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Touch   term used to refer to the sensations caused by mechanical displacements of the skin. These displacements occur any time you grasp, wield, or make contact w/an object  
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Tactile (adj. for touch)   refer to mechanical interactions when talking about touch and encompassing perception of temp. changes (thermal sensation), itchiness, & internal sensations (body in space)  
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Eyes & ears can   perceive signals frm objects that are far frm the body, but we must almost always be in direct contact w/an object to perceive it by touch  
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Human sense of touch   located in the skin  
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How touch receptors function   1. Type of stimulation (s/a diff. stimulus events) to which the receptor responds; 2. Size of RF; size of RF refers to the extent of the body area that elicits receptor response; 3. Rate of adaptation (fast vs. slow)  
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Fast-adapting receptors (FA 1 & 2)   responds to bursts of APs, 1st when its preferred stimulus its preferred stimulus is applied & then again. Doesn’t respond during the steady state b/w stimulus onset & offset  
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Slow-adapting receptors (SA 1&2)   remains active throughout the period during which the stimulus is in contact w/its receptive field (RF)  
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Tactile receptor   consists of a nerve fiber & an assoc. expanded ending. These receptors fall under the category of A-beta fibers: wide diameters that permit very fast neural conduction  
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Meissner corpuscle   located @ the junction of the epidermis & dermis; and is FA type 1  
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Merkel cell neurite complex   located @ the junction of the epidermis & dermis; and is SA type 1  
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Pacinian corpuscle   embedded in the dermis and subcutaneous tissue; and is FA type 2  
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Ruffini endings   embedded in the dermis & subcutaneous tissue; & is SA type 2  
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SA 1   sustained pressure, very low frequency (<~5Hz); texture perception. Also spatial deformation which is helpful in pattern/form detection  
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FA 1   temporal changes in skin deformation (~5-50 Hz); low frequency vibration detection  
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FA 2   temporal changes in skin deformation (~50-700 Hz); high frequency vibration detection  
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SA 2   sustained downward pressure, lateral skin stretch, skin slip (low sensitivity to vibration across frequencies); finger position & stable grasp  
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Thermoreceptors   normal internal body temp. is b/w 30-36 degrees Celsius. Also respond to objects in the environment and warm or cold fibers fire accordingly  
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Nocioreceptors   have bare nerve endings & respond to various forms of tissue damage or to stimuli that have the potential to damage tissue (incl. extreme skin temps. Lower than 15 degrees or higher than 45 degrees Celsius). A- delta fibers & C fibers  
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Painful events occur in 2 stages   (1) quick sharp burst of pain (2) throbbing sensation; reflect the onset of signals 1st frm A-delta fibers & then frm C-fibers  
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Spinothalamic pathway   slower; carries most of info. frm thermoreceptors & nociceptors. A lot of synapses w/in spinal cord which slows conduction while providing a mechanism for inhibiting pain pain perception when necessary  
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Dorsal column-medial lemniscal pathway (DCML)   incl. wider diameter axons & fewer synapses & conveys info. more quickly to the brain. Tactile & kinesthetic info. carried through this pathway; used for planning & executing rapid movements when quick feedback is a must  
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Phantom limb; Vilayanur Ramachandran’s research   amputees often feel sensations in their phantoms arms & hands when their faces or remaining limbs are touched. Idiosyncrasy in homunculus  
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Hand & arms areas of S1“invaded” by neurons carrying info. frm touch receptors in the face   Other parts of the brain listening to hand & arms areas are not fully aware of altered connections & therefore attribute activity in these areas to the stimulation frm the missing limb  
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“What” system   the ability to recognize objects through touch  
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“Where”   detection of spatial orientation/deformation abilities  
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Gate control theory   bottom-up pain signals frm nociceptors can be blocked via a feedback circuit located in dorsal horn. As gate neurons send excitatory signals sensory info. is allowed to go in but inhibitory signals frm the gate neurons cancel transmission to the brain  
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ACC   processes the raw sensory data frm S1 and S2 in such a way as to produce an emotional response  
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Secondary pain affect   emotional response assoc. w/long-term suffering that occurs when painful events are imagined or remembered  
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