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Landscape Ecology 1

Landscape Ecology Lecture Test One

QuestionAnswer
Landscape Ecology study of the causes and consequences of spatial heterogeneity across a range of scales
Carl Troll German scientist to introduce concept of landscape ecology based upon studies geography, vegetation sciences and aerial photography
Landscape Ecology study of landforms and the organisms which inhabit the area; considers the causes and consequences of habitats from a small scale to larger scales
Configuration specific arrangement of spatial elements; also referred as spatial structure or patch structure
Connectivity spatial continuity of a habitat or cover across a landscape
Corridor relatively narrow strip of particular type that differs from the areas adjacent on both sides
Cover Type category within a classification scheme defined by the user that distinguishes among the different habitats, ecosystems, or vegetation on a landscape
Edge Portion of an ecosystem or cover type near its perimeter and within which environmental conditions may differ from the interior location in the ecosystem; also used as a measure of the length of adjacency between cover types in a landscape
Fragmentation breaking up of a habitat or cover type into smaller, disconnected parcels
Heterogeneity quality or state of consisting of dissimilar elements, as with mixed habitats or cover types occuring on a landscape
Landscape area that is spatially heterogeneous in at least one factor of interest
Matrix background cover type on a landscape, characterized by extensive cover and high connectivity; not all landscapes have a definite matrix
Patch surface area that differs from its surroundings in nature or appearance
Scale measured by 2 factors : grain or extent; spatial or temporal dimension of an object or process
Factors for Emergence Landscape Ecology 1. Broad scale environmental issues and land management problems 2. Development of new-scale related concepts in ecology 3. Technological advances
Development of Landscape Ecology Resource management needs led to development to manage native plants and animals as land use and climates changed
Government Agencies Involved in Environmental Agencies Forest Service, Bureau of Land Management, National Park Services
Scale scope of study changes depending on the type of ecological problems; insight at one scale does not translate directly to another scale
Whitaker Gradient Analysis studied plant communities in Great Smokey Mountains and distinct vegetation patterns emerge with changes in elevation due to temperature and moisture(1952, 1956)
MacArthur & Wilson Island Biogeography -1. probability of a species reaching an island is inversely proportional to the distance from mainland and is directly proportional to island size 2. Probability of extinction on an island is function of island size
MacArthur & Wilson studied island biogeography which studied correlation between island size and distance from mainland to determine immigration and extinction of organisms
Equilibrium Theory of Island Biogeography MacArthur & Wilson study shows islands that are large and close have a lower rate of extinction and higher rate of immigration while small islands far away from the mainland have a lower rate of immigration and higher rate of extinction
Simberloff Island Biogeography study offers a good null model important for the design of nature reserves; study compares single, large habitats to several,small habitats [SLOSS}(1974)
Hanski Metapopulation Theory(1998)collection of interconnected subpopulation with source surrounded by areas known as sink, areas
Source area has higher fitness or capability to pass DNA to next generation
US Endangered Species Act Established 1.Recovery Plan 2. Critical Habitat Designation
Huffaker Experiments with frugivorous & predatory mites to study spatial manipulation of oranges shift dynamics between unstable(close together) and stable (Close together)
Huffaker Study showing spatial pattern can effect both stability of populations and total population size.
Spatial Patterns Result of complex interplay of abiotic factors, biotic factors and disturbances
Abiotic Factors environmental factors such as weather,dissolved oxygen, silt, temperature or elevation ; most important landscapes variable are climate and landform
Biotic Factors competition,predation,mutualism which effect populations which then effect communities
Disturbancces Examples include : clear cuts, burns, glaciers, hurricanes: effect abiotic and biotic factors and directly effects populations and communities
Population One species in certain area
Communities collection of organisms in a defined area
Population Dynamics Factors affecting include : diet, habitat use, population size, predator, reproduction including clutch size and clutch frequency. and sex ratio
Community Dynamics Factors affecting include: number of species, biomass, biological indices concerning condition of community such as IBI(fish), B-IBI (benthic invertebrates) QHEI(habitat index)
Broad Spatial Scale Understanding Needed For: Pattern and Processes Issues such as acid rain, global climate change, habitat fragmentation and conservation of biodiversity
Biotic Hierarchy Levels of organization (larger to smaller) include continental,terrestrial/aquatic, biomes, ecoregions,habitat ecotypes, trees, organisms living in trees...most meaningful interactions occurring at the lowest level or smallest scale
Hunter research studied avain community patterns which were dependent on spatial, temporal and taxonomic resolution. Songbirds require small spatial habitats while raptorial birds require continuous spatial habitats
Spatial Pattern Pattern inherent at one scale may disappear when you scale up because data is not always transferable.
Absolute Scale actual distance, direction, shape and geometry
Cartographic Scale degree of spatial reduction indicating the length used to represent a larger unit of measurement; ratio of distance on the map to distance on the earth's surface represented by the map, usually 1:10,000
Cartographic Scale large scale -> fine resolution small scale -> coarse resolution
Critical Threshold point at which there is an abrupt change in quality, property or phenomenon
Extent size of the study area or the duration of time under consideration
Extrapolate to infer from known values ; to estimate a value from conditions of the argument not used in the process of estimation
Extrapolate to transform information 1. from one scale to another(either grain size or extent 2. from one system(or data set) to another system at the same scale
Grain finest level of spatial resolution possible within a given data set
Hierarchy system of interconnections or organization where in higher levels constrain and control the lower levels constrain and control the lower levels to various degrees depending on the time constraints of the behavior
Holon representation of an entity as a two-way window through which the environment influences the parts and parts communicate as a unit to the rest of the universe (Koestler,1967)
Levels of Organization place within a biotic hierarchy
Relative Scale transformation of absolute scale to a scale that describes the relative distance, direction or geometry based on some functional relationship
Resolution precision of measurement; grain size if spatial
Scale spatial or temporal dimension of an object of process, characterized by grain and extent
Schoener (1976) documented an increase in area yields an increase in species richness; quantitatively shown with species richness equlaed to fitted constant multiplied by the area squared
Schoener species-area relationship is one of community ecology's few laws and tend to give asymptotic,with limit,curve
Preston Small,isolated island have fewer species per unit of area and higher slope values; small island are small targets and have higher probabilities of extinction
Founder Effect bredding within a population; loss of genetic variationdue topopulation started by a small number of individuals (special case of genetic drift)
Island Effects Gigantism - Giant Tortoise due to no predator and herbivore Dwarfism -Komodo Dragons - resource limitation
Haggert Scale Problems: 1. Scale Coverage Patterns 2. Scale Linkage Patterns 3. Scale Standardization Problem
Hierarchy Theory interconnected systems have higher levels which constrain lower. Ex. of this theory includes the local fish assemblage in a field site contain organisms remaining from filtering of upper levels along with abiotic factors, biotic factors and disturbances
Sampling Methods Transect - line count along a linear path Quadrat - count inside a square area
Scale Up most measurements in ecology made at transect or quadrat scale - need to extrapolate
King (1991) direct extrapolation uses data or model simulations froma number of samples in the landscape to estimate to larger areas
King (1991) extapolation by expected value involves imulation modeling
Schneider (1994) confidence intervals around a measurement made at one scale may not directly translate to another scale
Succession predictable change in vegetation communities immediately following a disturbance 1.primary succession completely destroys soil layer 2. secondary succession keeps soil intact
Categories of Spatial Patterns 1. Local Uniqueness ex: mountain 2. Phase Difference ex: succession 3. Dispersal ex: wind , water
Levin (1976) studied unique features of landscape such as uniqueness of locality, succession phase difference, and dispersal of plant species
Climate & Landforms Most important abiotic landscape variables which establish template for soils and biota
Biomes form primarily because of temperature and precipiation
Climate 1.warms earth's surface & energy used to fuel food webs 2. equator gets most sun 3. temperate latitudes have seasons 4. artic & antartic one day of total light and total darkness per year at solstice
Elevation effects temperature, density, pressure and moisture. Air density & pressure decrease as elevation increases.
Coriolis Effect deflects winds; winds blowing to equator deflect to west and known as trade winds and winds blowing to poles deflect east as westerlies
Landforms Influence Climate Examples: 1. coastal California lower temperature , inland Californai higher temperature 2. Death Valley, Nevada green,lush on west side mountain while east side of has desert conditiond
Milankovich Cycles climate cycles caused by the earth's wobbling on it axis; Glacial/Interglacial Cycles 90,000 years of gradual cooling followed by 10,000 years of rapid warming
Croin & Schneider (1990) Organisms response to climate change 1. Evolve & Speciate 2. Migrate Long Distances 3. Become Extinct
Mass Extinction Events ~6 events, largest being formation of Pangea because 1/2 shoreline which decreased habitat
Organism Response to Climate Change 1. Glacial-Interglacial cycles trigger disassembly of communities,followedby reassembly that can be unpredictable
Organism Response to Climate Change 2. past communities @ many sites feature mixtures of species that are absent or rare on modern landscape
Organism Response to Climate Change 3. displacement of entire vegetation zones or communities can occur
Swanson (1988) Effects of landform on ecosystem patterns & proceses: 1. Elevation,aspect,parent materials & slope of landforms affect air & ground temperature and quantities of moisture, nutrients, & other materials available
Swanson 2. Landforms affect the flow of organisms (wind) 3. Landforms affect frequency & spatial pattern of natural disturbance (fire, wind, grazing)
Swanson 4. Landforms constrain spatial pattern and rate or frequency of geomorphic processess (landslides or shifts to river channels)
Gause (1934) determined no two species can occupy the same niche by studying two different types of parmecium
niche everything an individual species needs to survive and reproduce ; Hutchinson defined as (n)dimensional hypervolume
Sutherland (1974) multiple states can occur when several species can potentially occupy and dominate site; can results in small stochastic(random) changes in initial conditions
Paine (1974) studied interactions in rocky intertidal zone which determined sea stars were keystone species that kept mussels in check; removal of sea stars caused increase in mussels which led decline in sea otters over competition for food source
Keystone Species keeps dominant species in check; doesn't allow an organism to overpopulate ; aka predator-mediated coexistence
Dominant Organisms 1. Define spatial patterns on landscape 2. Alter abiotic conditions & provide resource base & substrate for other populations in ecosystem 3. Rest of ecosystem is constrained to operate in spatial pattern (plants/terrestrial, coral/marine)
Dominant Organisms Examples: Redwood trees, American Bison
Human Land Use dramatically altered biodiversity of organisms 1. pollen & fossil record show shifts in patterns 2. Advent of agri. crops such as corn, wheat,and rice shifted land use patterns 3. Weeding & Nutrient Addition for permanent fields
Human Land Use 4. Loss of native diversity in Middle East and Eutope from long-term use 5. Native Americans in North America influenced landscape by settlements, agriculture, hunting & fire use 6. Agri,livestock,forest harvest,& construction have greatest threat
Dsts Used in Landscape Analysis 1. aerial photography 2. digital remote sensing 3. published data and censuses 4.field mapped data
Methods of Representing Spatial Data in GIS 1. Raster (celled) based 2. Vector based
Anderson (1976) Developed heirarchical land classification system used for landscape analysis; divided into 2 levels
Sources of Potential Error in GIS Data 1. Obvious Sources - age of data,aerial coverage,map scale,political boundaries 2. Natural Variation in Original Measurements- positional accuracy,content accuracy,variation in data sources
Sources of Potential Error in GIS Data 3. Processing Errors - numerical computation,topological analysis,classification
Problems for Landscape Pattern Analysis 1.Identification of Classification Scheme 2.Definition of Scale 3.Identification of Patch 4.Correlation of Metrics 5.Idenitification of Significant Change
Considerations to Decrease Error 1. Dificulty to specify biases for map of different extent 2.no single map extent optimal for all snalyses 3.Sample maps with greatest extent unless optimal sample size is known
O'Neill (1996) Proposed general rule to avoid bias in calculating landscape metrics; 1. Grain 2X-5X smaller than spatial feature 2. Extent 2X-5X larger than largest patches
Ritters Identified 5 independent landscape metrics 1.# of classes of cover types 2.texture, fine or coarse 3.patches,compact or dissected 4.patches,linear or planar 5.patch perimeter,complicated or simple
Landscape Metric Categories 1. Landscape Composition 2. Spatial Configuration 3. Fractals
Metrics of Landscape Composition 1.Fraction or Proportion 2.Relative Richness 3.Diversity & Dominance 4.Connectivity
Metrics of Spatial Configuration 1.Probabilities of Adjacency 2.Contagion 3.Patch Area & Perimeter 4.Connectivity 5.Proximity Index 6.Area-Weighted Average Patch Size
Fractals 1.Fractal Dimension 2.Fractal as Measure of Patch Shape
Selection of Multiple Metrics Subset should 1.explain pattern variability without redundancy 2.capture relevent aspects of pattern to answer to study question 3.use multivariate statistics
Ritters Recommendation of Multivariate Measures for Landscape Analysis to Avoid Redundancy 1.total # of land cover types 2.contagion 3.fractal dimension 4.average patch perimeter-area ratio 5.relative patch area
McGarigal & Marks (1995) Recommendation of Multivariate Measures for Landscape Analysis to Avoid Redundancy 1.patch shape & edge contrast 2.patch density 3.patch size
Li & Reynolds (1994,1995) Theoretical considerations to compute aspects of spatial heterogeneity 1.#of land-cover types 2.proportion of each type on the landscape 3.spatial arrangement of patches 4.patch shape 5.contrast between neighboring patches
Purpose of Spatial Statistics 1.identify the spatial scales over which patterns or processes remain constant 2.to interpolate or extrapolate point data to infer the spatial distributions of variables of interest
Important Question Addressed by Spatial Statistics 1. What is the appropriate scale to conduct an anlysis? 2.What is the nature of spatial structure of a particular variable?
Correlograms in analysis of time series,commonly-used tool for checking randomness in a data set
Semivariograms defines spatial scales over which patterns are dependent ; are calculated by rearranging the data into N data pairs separated by distance ; flat variograms indicates patterns that lacks spatial structure
Criteria for Metrics to Quantify Landscape Patterns 1. Selected to paticular question or objective 2. Distributed over full range of potential values & behavior of metrics known 3.indexes ahould be relativey independent of each other
Spatial Statistics quantify aspects of landscape patterns and used to detect the spatial scales of autocorrelation for landscape elements or to interpolate point data to infer spatial distributions of a variable of interest
Potential Areas of Research Development for Landscape Pattern Analysis 1. Statistical Properities & Behavior of Metrics 2. Relative Sensitivities of Different Metrics to Detect Changes in Landscape 3.Documentation of Empirical Relationship between Landscape Patterns and Ecological Processes of Interest
Proportion metric of landscape composition used to calculate the amount of landscape that is occupied by each cover type
Relative Richness metrics of landscape composition which calculates the number of cover types present, without regard to spatial arrangement,as a percent of total number of possible cover types
Diversity measure of landscape composition which measures how evenly the proportions of cover types are distributed; relative evenness; high value = high evenness low value = low evenness
Dominance metric of landscape composition which is the deviation from the maximum possible diversity high value=one or few cover types low value=cover types in close proportion
Connectivity metric of landscape composition which is a pattern across a landscape represented by a series of nodes and linkages;known as gamma index low gamma index=low connectivity high gamma index=high connectivity
Probability of Adjacency measure of spatial configuration which measures the probability that a grid cell of one type is adjacent to grid cell of another type; can be computed directionally
Contagion measure of spatial configuation distinguishes between overall landscape patterns that are clumped or dissected; high value=high clump pattern of cover low value=dsipersed cover
Patch Area & Perimeter measure of spatial configuration used to show frequency distribution of numbers of patches by patch size,cumulative frequency distribution of patch sizes,simple mean, standard deviation of patch size or area-weighted mean patch size
Patch Area & Perimeter high value=complex boundary low value=simple boundary
Connectivity measure of spatial configuration which measures the relative size of the largest patch of habitat or the inverse,fragmentation, to calculate the average distance between patche
Proximity Index measure of spatial configuration which measures the degree to which patches in the landscape are isolated fromother patches; high value=connected patches low values=isolated patches
Area-Weighted Average Patch Size measure of spatial configuration used to reflect the expected patch size that would be encountered by simple random placement of points on the map
Fractal Dimension measure which shows the level of variation present at all scales
Mandelbrot (1985) defined fractal as a shape made of parts similar to the whole in some way
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