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BIOL1202 exam3

chapters 28 thru 33

TermDefinition
clade Excavata (monophyletic) · characterized by cytoskeleton, some have excavated feeding groove on one side of body · three monophyletic groups: diplomonads, parabasilids, euglenozoans
Excavata: diplomonads and parabasilids · lack plastids and have modified mitochondria, most anaerobic
Excavata: diplomonads · reduced mitochondria - mitosomes, get energy from anaerobic biochemical pathways · two equal-sized nuclei, many flagella · often parasites (Giardia intestinalis)
Excavata: parabasilids · reduced mitochondria - hydrogenosomes, get some energy anaerobically · includes ST parasite Trichomonas vaginalis
Excavata: euglenozoans · includes predatory heterotrophs, photosynthetic autotrophs, mixotrophs, parasites · specifically - kinetoplastids and euglenids · main feature: spiral, crystalline rod within flagella
Excavata: euglenozoans - kinetoplastids · single mitochondrion with organized mass of DNA - kinetoplast · free-living species consume prokaryotes in nearly every environment · trypanosomes: parasitic on plants, animals, other protists, ex. African sleeping sickness, Chagas' disease
Excavata: euglenozoans - euglenids · one or two flagella emerging from pocket at end of cell · some are mixotrophs - can be autotrophic or heteroptrophic depending on environment
supergroup SAR (monophyletic) · stramenopiles, alveolates, rhizarians · controversial supergroup
SAR: stramenopiles · some of the most important photosynthetic organisms on Earth - diatoms, golden algae, brown algae · most have one "hairy" flagellum paired with one smooth one
SAR: stramenopiles - diatoms · unicellular algae with a two-part, glass-like cell wall of silicon dioxide · highly diverse, major component of phytoplankton · oceanic diatom bloom -> dead individuals bring atmospheric carbon to ocean floor
SAR: stramenopiles - brown algae · largest and most complex algae, all multicellular, most marine, commonly called seaweeds (similar to plants but lack true tissues) · rootlike holdfast, stemlike stipe, leaflike blades, some have floats to keep photosynthetic components near surface
SAR: alveolates · membrane-enclosed alveoli just under plasma membrane · includes dinoflagellates, apicomplexans, ciliates
SAR: alveolates - dinoflagellates · two flagella, each cell reinforced by cellulose plates · abundant components of marine and freshwater phytoplankton · phototrophs, mixotrophs, and heterotrophs · cause toxic "red tides"
SAR: alveolates - apicomplexans · most parasites of animals, some cause human disease - infectious sporozoites penetrate host cells · most have sexual and asexual life stages and need 2+ hosts · Plasmodium parasite requires mosquito and human host to complete life cycle - malaria
SAR: alveolates - ciliates · use of cilia to move and feed, two types of nucleus, most prey on bacteria or other protists, ex. Paramecium · genetic variation from conjugation - two individuals exchange haploid micronuclei (sexual process) · reproduction - usually binary fission
SAR: rhizarians · include radiolarians, forams · many species are amoebas - move and feed using pseudopodia - extensions of cell surface · differ from amoebas in other clades by threadlike pseudopodia
SAR: rhizarians - radiolarians · delicate, symmetrical internal silica skeleton · pseudopodia reinforced by microtubules radiate from central body, cytoplasm covering microtubules engulf prey stuck on pseudopodia
SAR: rhizarians - foraminiferans · porous shells called tests made of calcium carbonate · pseudopodia extend thru pores in test, some house photosynthetic algae within test · freshwater and marine · magnesium content in fossils used to estimate ocean temp. changes over time
supergroup Archaeplastida · includes red algae, green algae, plants
Archaeplastida: red algae · accessory pigment phycoerythrin masks green chlorophyll -> red color, varying from red in shallow water to almost black in deep water · usually multicellular, largest are seaweeds · most abundant large algae in tropical waters - farmed for carageenan
Archaeplastida: green algae · grass-green chloroplasts, closely related to plants, most freshwater but many marine and some terrestrial · paraphyletic group: charophytes (closer to true plants) and chlorophytes · forms colonies and true multicellular bodies -> large size
supergroup Unikonta · includes animals, fungi, some protists · two clades: amoebozoans and opisthokonts · unclear if unikonts separated from other eukaryotes earlier or later
Unikonta: amoebozoans · amoebas with lobe/tube-shaped pseudopodia · includes slime molds, tubulinids, entamoebas
Unikonta: amoebozoans - slime molds · resembles fungi due to convergent evolution · plasmodial: unicellular feeding mass (plasmodium) · cellular: motile stage, forms fruiting bodies
Unikonta: amoebozoans - tubulinids · common in soil, freshwater, marine · most are heterotrophic and actively seek and consume bacteria and other protists
Unikonta: amoebozoans - entamoebas · parasites of vertebrates and some invertebrates · Entamoeba histolytica causes amoebic dysentery
shared traits between plants and charophytes · rings of cellulose-synthesizing proteins · structure of flagellated sperm · formation of phragmoplast
traits in plants but not charophytes · alternation of generations · multicellular, dependent embryos · sporangia makes walled spores · multicellular gametangia · apical meristems
gametophyte vs. sporophyte · gametophyte: haploid (n), produces haploid gametes thru mitosis · sporophyte: diploid (2n), makes haploid spores thru meiosis · spores give rise to gametophytes
archegonia vs. antheridia · archegonia: female gametangia, produces a single, non-motile egg · antheridia: male gametangia, produces and releases sperm · each egg is fertilized within archegonium
bryophytes · nonvascular plant · three phyla of small, herbaceous plant: hepatophyta (liverworts), bryophyta (mosses), anthocerophyta (hornworts) · earliest lineages to diverge from common ancestor of land plants
bryophytes - gametophyte · dominant generation, but height constrained by lack of vascular tissue · rhizoids anchor them to substrate, produce flagellated sperm in antheridia and egg in each archegonium · may reporuduce asexually in time of drought
bryophytes - sporophyte · never live independently from gametophyte · smallest and simplest sporophyte of all plants · consists of foot, seta (stalk), and sporangium (aka capsule) which makes spores
vascular plants · life cycle with dominant sporophytes · vascular xylem and phloem · well-developed roots and leaves · spore-bearing leaves - sporophylls
vascular plants - xylem · vascular tissue that conducts most of the water/minerals and has tube-shaped cells - tracheids · cells strengthened by lignin (structural support)
vascular plants - phloem · vascular tissue with cells arranged into tubes that distribute sugars, amino acids, other organic products
vascular plants - leaves · microphylls: small, single vein · megaphylls: large, branched vasculature
vascular plants - sporophylls and spore variations · sporophylls: modified leaves and sporangia · sori: sporangia clusters on sporophyll undersides · strobili: cone-like groups of sporophylls · homosporous species -> one spore type -> bisexual gametophyte
vascular plants - sporophylls and spore variations · heterosporous species -> megaspores -> female gametophytes · microspore -> male gametophyte
phylum Lycophyta: club mosses, spike mosses, quillworts · small herbaceous plants · has vascular tissue, not true moss
phylum Monilophyta: ferns, horsetails, whisk ferns and relatives · ferns are most widespread seedless vascular plants - more than 12,000 species · most diverse in tropics but also thrive in temperate forests
seed plants - seed · consists of embryo, nutrients surrounded by protective coat
seed plant characteristics · reduced gametophytes · heterospory · ovules (eggs) · pollen
seed plants - advantages of reduced gametophytes · microscopic gametophytes develop in walls of spores retained within parent sporophyte - protects developing gametophyte from environmental stress and gets nutrients from parent sporophyte
seed plants - heterospory · megasporangia produce megaspores -> female gametophyte · microsporangia produce microspores -> male gametophyte
seed plants - ovule · consists of megasporangium, megaspore, and 1+ protective integuments · gymnosperms usually have one integument, angiosperm usually two
seed plants - pollen · microspore develops into pollen grain - male gametophyte in pollen wall · germinated pollen grain -> pollen tube -> sperm to female gametophyte in ovule · pollen does not require water to travel
seed evolutionary advantages · can remain dormant until conditions are good for germination · has food supply · can be transported long distances via wind or animals
gymnosperms · most are cone-bearing conifers · mini gametophytes, produces dispersible seeds, transfers sperm to ovules thru pollen · male pollen cones: small, modified leaves (microsporophylls) · female ovulate cones: larger, both modified leaf and stem tissue
gymnosperms: phylum Cycadophyta · large cones, pine-like leaves · unlike most seed plants, has flagellated sperm · thrived during Mesozoic but most of few surviving sp. are endangered
gymnosperms: phylum Ginkgophyta · single species: Ginkgo biloba · like cycads, also has flagellated sperm high tolerance to air pollution, popular ornamental tree and supplement
gymnosperms: phylum Gnetophyta · three genera: Gnetum, Ephedra (diet aid), Welwitschia · species vary in appearance, some tropical, others desert
gymnosperms: phylum Coniferophyta · largest of all gymnosperms · most have woody cones, few have fleshy cones · most are evergreens and photosynthesize year-round · many commercial uses (flavoring gin)
angiosperms · seed plants with reproductive structures - flowers and fruits · most widespread and diverse plants · single phylum - Anthophyta
angiosperms - flowers · specialized for sexual reproduction · use insects, animals, wind, to pollinate · a specialized shoot with up to four types of modified leaves - flower organs · variable in shape/symmetry, size, color, odor
angiosperms - flower organs (non-repro) · sepals: enclose and protect flower · petals: often brightly colored to attract pollinators unless wind is used
angiosperms - flower organs (repro) · stamens: male repro organs, stalk called filament and sac called anther (microspore -> pollen) · carpels (pistil): female repro organs, ovary at base of style with sticky stigma at tip to receive pollen · ovary has ovule -> fertilization -> seed
angiosperms - fruits · formed when ovary wall thickens and matures · protect seeds, aid in dispersal (wind, water, animals), may be fleshy or dry
angiosperm life cycle · flower has both male and female structures - anthers have microsporangia (makes pollen), stigma has ovary at base (embryo sac within ovule) · pollen grain lands on stigma -> germination -> pollen tube of male gametophyte grows down to ovary
angiosperm diversity · monocots: one cotyledon · dicots: two cotyledons · eudicots: most dicots · basal angiosperms: oldest lineages · magnoliids: share traits with basals but evolved later
angiosperms - basal angiosperms · ~100 sp., three small lineages · water lilies, star anise, Amborella thrichopods
angiosperms - magnoliids · woody or herbaceous · more closely related to monocots and eudicots than basal angiosperms
angiosperms - monocots · about 1/4 of angiosperms, ~70,000 sp. · largest groups are orchids, grasses, palms
angiosperms - eudicots · more than 2/3 of angiosperms, ~170,000 sp. · includes legumes, roses
key traits of fungi: heterotrophs · use enzymes to break down absorbed complex molecules -> smaller organic compounds · decomposers: nutrients from non-living organic material · parasites: nutrients from live host · mutualists: nutrients from live host, reciprocates with host benefit
key traits of fungi: body structure · most common body structures: multicellular filaments and single cells (yeasts) · body of filamentous fungi forms branched hyphae adapted for absorption with tubular cell wall with chitin
key traits of fungi: hyphae · most have hyphae divided by septa, with pores for cell-cell organelle mvmt · coenocytic fungi lack septa, have continuous cytoplasmic mass with endless nuclei · hyphae form interwoven mass - mycelium, maximizes surface-volume ratio -> efficiency
fungi: specialized hyphae · some are specialized for feeding on live animals · haustoria: extracts plant nutrients · arbuscules: used by mutualistic fungi to exchange nutrients with plant hosts
fungi: mycorrhizae · mutually beneficial relationships between fungi and plant roots · deliver phosphate ions, minerals, ammonium to plants · most vascular plants have mycorrhizae, which colonize soils by dispersal of haploid spores
fungi: sexual reproduction · fungal nuclei are normally haploid; hyphae can fuse from two different mating types and use pheromones to communicate type · plasmogamy (union of cytoplasm ) long time before karyogamy(nuclear fusion) -> diploid cells -> meiosis (genetic variation)
fungi: asexual reproduction · molds produce haploid spores by mitosis and form visible mycelia · yeasts reproduce by budding; many molds and yeasts have no known sexual stage (deuteromycetes)
Opisthokont clade · fungi, animals, protistan relatives · evolved from unicellular flagellated ancestor
phylum Chytridiomycota (chytrids - water molds) · nearly every habitat including hydrothermal vents · decomposers, parasites, mutualists, most likely diverged early in fungal evolution · unique with zoospores - flagellated spores
phylum Zygomycota (zygomycetes) · fast-growing molds, parasites, commensal symbionts · coenocytic septa, asexual sporangia produce haploid spores · named for sexual zygosporangia (when hyphae touch) - site of karyogamy then meiosis, survive unfavorable conditions
phylum Glomeromycota (glomeromycetes) · nearly all species form arbuscular mycorrhizae
phylum Ascomycota (ascomycetes) · nearly all habitats, include plant pathogens, decomposers, symbionts · produce sexual spores in asci in ascocarps - fruiting body, also repro asexually · "sac fungi" ranging in size from unicellular to elaborate cup fungi and morels, >25% form lichens
phylum Basidiomycota (basidiomycetes) · includes common mushroom, puffball, shelf fungi · some form mycorrhizae, others are plant parasites, many wood decomposers · defined by transient diploid clublike life stage - basidium, also long-lived dikaryotic mycelium
fungi as mutualists ·relationships with plants, algae, cyanobacteria, animals · absorb nutrients from host -> host benefit · very important in natural ecosystems and agriculture · plants harbor harmless fungi endophytes within leaves or other parts - usually ascomycetes
fungus-animal mutualisms · digestive services - fungi help break down plant material in guts of cows, other grazers · many ant species keep fungi in "farms" to aid in digestion
fungi: lichens · symbiotic association with photosynthetic microorganism (green algae or cyanobacteria) and fungus (usually ascomycete) · algae/cyanobacteria occupy inner layer between mass of fungal hyphae ·important on new rock/soil surfaces
fungi as parasites · around 30% of fungal species are parasites/pathogens, usually of plants - some toxic to humans (ergotism) · animals much less susceptible than plants but chytrid B. dendrobatidis attacks amphibian species
fungi infecting humans · mycosis: human fungal infection · ringworm, athletes foot · systemic mycoses spred thru body - ex. Coccidioidomycosis -> tuberculosis-like symptoms · some opportunistic - Candida albicans -> yeast infection
practical uses of fungi · humans eat fungi and use others to make cheeses, alcohols, breads ·some used to make antibiotics (ascomycete Penicillum) · genetic research -> applications in biotechnology - yeast insulin-like growth factor for diabetes
animal characteristics · heterotrophs that ingest food · multicellular eukaryotes · cells supported by structural proteins like collagen rather than cell walls · nervous and muscle tissues
animal reproduction and development · sexual diploid stage dominates life cycle · zygote undergoes rapid cleavage after fertilization -> hollow blastula - gastrulation forms gastrula with 3D layers of tissue · most have at least one immature larval form · Hox genes regulate body form
fossils from Cambrian explosion characteristics · bilaterally symmetrical · complete digestive tract · one-way digestive system
hypotheses for Cambrian explosion and Ediacaran decline · new predator-prey relationships · rise in atmospheric oxygen · evolution of Hix gene complex, new microRNAs
Neopreoterozoic era (1 BYA - 541 MYA) · early members of animal fossil record - Ediacaran biota dates back 560 MY · microscopic animal embryos found in Neoproterzoic rocks · evidence of predation
Paleozoic era (541 - 252 MYA) · Cambrian explosion (535-525 MYA) is earliest appearance of major groups of living animals
Mesozoic era (252 - 66 MYA) · coral reefs emerge - important marine ecological niches · ancestors of plesiosaurs - reptiles that returned to oceans · dinosaurs are dominant terrestrial vertebrates, mammals emerged, flowering plants and insects diversified
Cenozoic era (66 MYA - present) · followed terrestrial and marine mass extinctions - large, non-flying dinosaurs and marine reptiles · mammals increased in size, explored vacated ecological niches · global cooling
radial symmetry · think of a flowerpot · animals have a top/bottom but no left/right · animals often sessile (anchored) or planktonic (drifting/weakly swimming)
bilateral symmetry · think of a shovel · animals have dorsal/ventral sides, left/right sides, anterior/posterior ends · many have sensory equipment (ex. brain) concentrated anterior · animals usually move actively, have central nervous system
tissues · grouped specialized cells isolated from other tissues by membranous layers · three germ layers give rise to tissues and organs of animal embryo in development · ectoderm covers embryo's surface, endoderm lines developing digestive tube (archenteron)
tissues - development · diploblastic animals have only ecto and endoderm layers (cnidarians and few others) · triploblastic animals have intermediate mesoderm (all bilaterally symmetrical animals)
body cavities · true body cavity called coelom - derived from mesoderm · coelomates are animals with true coelom · pseudocoelomates have body cavity from meso and endoderm (endo uncovered) · acoelomates lack a body cavity
body cavity functions · fluid cushions suspended organs · fluid acts as skeleton · cavity enables internal organs to grow/move independently from body wall
development - cleavage · protostomes: cleavage spiral and determinate (cell fate already known) · deuterostomes: cleavage radial and indeterminate (each cell has capacity to become full embryo - twins and embryonic stem cells)
development - coelom formation · protostomes: splitting pf solid masses of mesoderm -> coelom · deuterostomes: mesoderm buds from wall of archenteron -> coelom
development - blastopore · blastopore forms during gastrulation and connects archenteron to exterior of gastrula · protostomes: blastopore -> mouth · deuterostomes: blastopore -> anus
bases for animal phylogenies · whole-genome analysis · morphological traits · rRNA genes · Hox genes · protein-coding nuclear genes and mitochondrial genes
phylogeny -> relationships among living animals · all animals share common ancestor · sponges are sister group to all other animals · clade Eumetozoa: "true animals", has tissues (not sponges) · most animal phyla are in clade Bilateria -> three major clades, all inverts, except Chordata
bilaterian clades · deuterostomia, ecdysozoa, lophotrochozoa · deuterostomia: hemichordates (acorn worms), echinoderms (sea stars + relatives), chordates · ecdysozoa: external skeletons, shed thru ecdysis · lophotrochozoa: lophophore to feed, distinct throchophore larva
phylum Porifera · sedentary, fresh and salt water, lack tissues, filter feeders thru cavity called spongocoel · several cell types (flagellated choanocytes, noncellular mesohyl layer, totipotent amoebocytes · most hermaphrodites
clade Eumetozoa · all animals except sponges and few other groups - true tissues
phylum Cnidaria · jellies, corals, hydra · single opening for mouth and anus, gastrovascular cavity · sessile polyp adheres to substrate, mobile medusa has bell-shaped body with mouth on underside · tentacles with cnidocytes - stinging cells · anthozoan - polyp only
phylum Platyhelminthes · flatworms - many parasites (flukes, tapeworms), acoelomates, gastrovascular cavity · dorsoventrally flattened body shape for best osmosis/diffusion
phylum Mollusca · snails and slugs, oysters and clams, octopi and squids · soft-bodied, most protected by calcium carbonate shell · similar body plans: muscular foot, soft/visceral mass, mantle · many have ciliated trocophore larval stage
phylum Annelida · coelomates with bodies made of series of fused rings · polychaetes: usually in ocean, feathery gills (gas exchange and filter feeding) · oligochaetes: earthworms, can fragment asexually · hirudinea: leeches, suck host's blood
Ecdysozoans - nematodes · found nealry everywhere · alimentary canal but no circulatory system, longitudinal body wall muscles -> thrashing motion · Trichinella spiralis - human parasite, can be acquired thru undercooked pork
Ecdysozoans - arthropods · segmented body, hard exoskeleton, jointed appendages · body plan dates back to Cambrian explosion · covered by cuticle - exoskeleton made of layers of protein and polysaccharide chitin
arthropods - chelicerates · clawlike feeding appendages · sea spiders, horseshoe crabs, ticks, mites, spiders
arthropods - myriapods · millipedes (detritivores) and centipedes (carnivores) - all terrestrial
clade Pancrustacea · insects and crustaceans together
Crustaceans · all environments, isopods, decapods
Hexapoda (insects and relatives) · every environment except rare in marine · wings - extension of cuticle · incomplete metamorphosis: nymphs resemble adults and molt to full size · complete metamorphosis: larva -> pupa -> adult
clade Deuterostomia · Echinoderms - sea stars and relatives · Chordates - vertebrates
Echinoderms · slow-moving or sessile, marine · thin epidermis covers endoskeleton (hard calcareous plates) · unique vascular system - hydraulic canals branching into tube feet for locomotion and feeding
phylum Chordata · two basal groups of inverts along with verts · bilaterally symnmetrical coelomates with segmented bodies · did not evolve from echinoderms, have evolved separately for ~500MY
Created by: junoregan
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