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Biology Exam #2

Chapters 32, 53, 55, 57, 59

QuestionAnswer
Two triploblastic clades? Protostomes and Deuterostomes
Which type is more common? Protostomes, which make up >94% of animals.
Major clades of protostomes? (2) Lophotrochozoans and Ecdysozoans
What do all protostome animals have? -three germ layers (triploblastic) -bilateral symmetry (at least part of life) -anterior brain near mouth -ventral nervous system w/ paired/fused nerve cords
Similar development of protostomes -spiral and determinate cleavage -Schizocoelous body cavity forms -mouth develops from blastopore
Ancestor of protostomes had a coelom but some diverged -Flatworm: acoelomate -Arthropods: have hemocoel (blood chamber) -Mollusks: open circulatory system but retained enclosed coelom
Lophotrochozoans (1st group of protostomes) -internal skeletons -many have trochophore larvae -some have a lophophore (circular/U-shaped feeding structure)
Ecdysozoans (2nd group of protostomes) -external skeleton -grows by ecdysis (molting) -all have single common ancestor
Arrow worms?? -similar to deuterostomes but identified as a protostome (due to molecular data) -possible sister group to all other protostomes -predators of plankton -approx. 100 species -2 pairs of lateral fins and tail fin
Molting -shedding exoskeletons that are replaced by new ones so the ecdysozoans can grow -evolved 500 million years ago
Cuticle -thin exoskeletons that cover some worm-like ecdysozoans -allows gas and water exchange (restricts them to moist places)
Arthropods (clade under ecdysozoans) -hard exoskeletons made of proteins and chitin(strong polysaccharide)
Bryozoans "moss animals" (group under lophotrochozoans) -colonial and secretes material that forms a "house" -in some species, individuals can become specialized -reproduce sexually -sperm released in & carried by water -individuals can rotate lophophore to increase contact w/ prey
Flatworms -acoelomates -approx. 25,000 species; most parasitic -include tapeworms and flukes -have cephalization, bilateral, and blind sac for stomach
Synapomorphy of phoronids and brachiopods? have lophophores that evolved independently from the bryozoans
Brachiopods -"lamp shells' -solitary marine animals -sessile and lack a head -larvae are planktonic -peaked in abundance in Paleozoic and Mesozoic (approx. 26,000) -only 355 species today
Annelids -segmented coelomates with specialization -includes polychaetes, oligochaetes, and leeches -thin permeable body wall for gas exchange -live in moist habitats -have bristles for crawling
Polychaetes -not a single clade -mostly marine worms -parapodia: extensions body wall & function in gas exchange, and movement -includes pogonophorans
Pogonophorans -deep sea tube worms that lost digestive tract -secrete tubes of chitin in which they live -live near hydrothermal vents -take up dissolved organic matter
Mollusks -most diverse group of lophotrochozoans -unsegmented coelomates -approx. 95,000 species -
4 Main groups of mollusks 1. Chitons 2. Bivalves (clams, oysters, scallops, mussels) 3. Gastropods (snails, sea slugs, slugs, limpets, abalones) 4. Cephalopods (squids, octopuses, nautiluses).
Parts of a mollusk -Foot: muscular structure, originally for locomotion and support of internal organs -Visceral mass: heart, digestive, excretory, and reproductive organs. -Mantle: fold of tissue that covers organs in the visceral mass & secretes the calcareous shell.
Open circulatory system of mollusks Blood and fluids empty into the hemocoel, where oxygen is delivered to internal organs
8 Phyla of Ecdysozoans (PKL H. NOTA) -priapulids -kinorhynchs -loriciferans -horsehair worms -nematodes -onychophorans -tardigrades -arthropods
Priapulids, Kinorhynchs, and Loriciferans -worm-like & marine -have thin cuticles that molt as they grow
Priapulids -unsegmented -three segments -burrow in soft sediments -capture prey with a pharynx
Kinorhynchs -microscopic -13 segments -feed by ingesting sediments through their retractable proboscis.
Loriciferans -small -body covered by six plates -"lorica": closet
Horsehair worms -very thin -up to 2 meters -mostly freshwater. -larvae are internal parasites of insects and crayfish. -adult has no mouth and a reduced gut.
Nematodes (round worms) -unsegmented -free-living & parasitic -approx. 25,000 species -thick multi-layer cuticle -parasites of humans cause trichinosis, filariasis, and elephantiasis -C. elegans
Onycophorans -closely are related to arthropods but lack specialization -Segmented worms -predators. -150 species; live in humid tropical areas. -thin, flexible cuticle -use fluid-filled body cavity as a hydrostatic skeleton for movement.
Tardigrades -water bears -same cavity of onycophorans -very small, no circulatory or gas exchange systems -when water lost can shrink to form a dormant state that can last at least a decade
Arthropods -most diverse animal (over 1 million species) -each segment has muscles that operate a segment and its appendages -complex movement patterns -Rigid exoskeleton provides support and safety
Trilobites -first appearance of jointed legs -flourished in Cambrian and Ordovician -went extinct at the end of the Permian
Myriapods & Chelicerates (groups of arthropods) have body with two regions: head and trunk
Myriapods -centipedes and millipedes -head with very long trunk & many legs -centipedes are carnivorous but millipedes are herbivores
Chelicerates -head has pair of appendages that act as mouthparts -includes pycnogonids (small sea spiders) and horseshoe crabs and arachnids
Horseshoe crabs -bottom-dwellers found in shallow water -go to intertidal zone to mate and lay eggs -changed very little
Arachnids -spiders, scorpions, mites, and ticks -have chelicerae, pedipalps, and 8 legs -“Silk” threads are produced by modified abdominal appendages
Crustaceans (still arthropods) -dominant marine arthropods -includes shrimp, lobster, crayfish, and crabs -most have head, thorax, and abdomen -many have a carapace
Carapace a fold of exoskeleton that extends over the head and thorax region
Hexapods (arthropods) i.e. insects -head, thorax, and abdomen -antennae on head & 3 legs on thorax -air sacs and tubular channels called tracheae extend from external openings (spiracles) into tissues throughout the body.
Wingless hexapods -includes Springtails, bristletails, and silverfish -wingless relatives of insects with internal mouth parts -probably resemble the insect ancestor
Pterygote insects -have two sets of wings -dragonflies can't hold wings against their body (ancestral)
Metamorphosis -substantial changes that occur between stages. -Incomplete metamorphosis: changes are gradual. -Complete metamorphosis: changes are drastic
Instars immature stages between molts
Neopterans -all other pterygote insects that can fold their wings -some have incomplete metamorphosis -Holometabolous insects have complete metamorphosis
When did insects and crustaceans separate? 450 million years ago (mya)
Factors that contribute to success of protostomes segmentation, complex life cycles, parasitism, diverse feeding structures, hard body coverings, and better locomotion
Chapter 53 :D
Ethology study of animal behavior from an evolutionary standpoint
Proximate mechanisms neuronal, hormonal, anatomical mechanisms
Ultimate causes selection pressures that shaped evolution of the behavior
Stereotypic behavior -always exactly the same -often specific to a species -i.e. specific web spinning by spider species
2 important experimental approaches Deprivation experiments and Genetic experiments
Deprivation experiments -young animals are reared with no experiences related to the behavior under study -i.e. squirrel in cage on liquid diet tried to bury nut when given one
Genetic experiments -provide insights into the genetic basis of behavior -selective breeding in plants & dogs
Genetic behavior in ducks -experiments done by Konrad Lorenz -males performed courtship displays specific to species -hybrids had elements of each "dance" & were't picked by females (sexual selection)
fosB in female mice female mice with fosB gene provide care for their pups; mutants with an inactive gene ignore their pups. -the fosB gene stimulates changes in the hypothalamus
Genetic control of behavior -can be adaptive (i.e. contribute to success of next generation) -In non-overlapping generations: offspring cannot learn behavior from parents
Releaser –an object, event, or condition required to elicit a behavior -i.e. the nut was a releaser for the squirrel
Critical period time in an animal's life where learning takes place
Imprinting animal learns a set of stimuli during a critical period, recognition of parents and offspring
Habitat Selection -Habitat must provide food, shelter, nest sites, escape routes -i.e. abalone veliger larvae recognize a chemical signal from corraline algae, their best food source, and settle there
Visual Cues -used for habitat selection -animals look at which animals are already present to determine if -i.e. collared flycatchers look into nests of other birds; brood size indicates good habitat quality
Territory area in which animals make their presence known and which excludes conspecifics (others of same species)
Cost-benefit approach -used to analyze behaviors -animal has limited energy for activities, animals cannot perform behaviors if energetic cost is greater than benefits of behavior
3 components of cost Energetic (energy difference expended if resting vs. performing in behavior), Risk (increased chance of being hurt/killed as a result of behavior), and Opportunity Cost (benefits the animal forfeits)
Foraging theory -uses cost-benefit analysis to study food choice -animals make choices among available prey in order to maximize the rate of energy intake -When food abundant, animal should take that and ignore rarer food
Circadian rhythm -daily cycle of activities -length: period -point: phase
Nocturnal vs. Diurnal -diurnal animals are more visual -nocturnal ones depend more on hearing, smell, touch, and have retinas made for low light
Photoperiod change in day length and indicator of season change
Circannual rhythms built-in neural calendars
Piloting knowing and remembering the structure of the environment (i.e. gray whales from Mexico to Bering Sea)
Homing ability to return to the same place from long distances (i.e. pigeons)
Migration movement of a whole species with the changes of season (i.e. birds returning to breeding grounds)
Bicoordinate (true) navigation –requires knowing current position relative to destination (i.e. albatrosses)
Birds in planetarium If the planetarium didn’t rotate while they were raised, the birds didn’t orient in any direction (birds may also sense magnetic fields)
Animal communication -can be signals or gestures -visual signals easy to produce, but not useful at night -loud vocalizations indicate strong males
Pheromones -chemical signals between individuals -can convey species, size, reproductive status, etc -can stay in environment for long time
Auditory signals -can be used at night, over long distances, and in complex environments -not as much info as visual
Tactile interactions -common among animals -ex. dance of the honeybees (round dance if food <80m away)
Social behavior happens when individuals gain greater fitness by working together than alone
Costs of social behavior -more competition for mating or getting food -increased risk of disease -
inclusive fitness an organism's individual fitness (# of gene copies of an individual in the next generation), plus the number of equivalents of its own offspring it can add to the population by supporting others.
Altruistic acts -behavior that may reduce helper’s fitness, but increases fitness of individual helped -naturally favored
Eusociality -highest level of organization of animal sociality -cooperative brood care -often have nests for large group -includes termites, ants, bees -usually a fertile queen and infertile workers
Benefits of social behavior -high population densities -greater defense
Chapter 55 >:P
Population dynamics patterns and processes of change in populations
Age structure age distribution of individuals, and how those individuals are spread over the environment
Population density # of individuals per unit area
Demographic events -deaths, births, emigration, immigration -create population dynamics
Full census possible when population is small and animal is large (i.e. elephant reservation in Kenya)
Molecular markers -used to track individuals -H isotopes in birds
Mark-recapture method -marking of some individuals, then capturing another sample of individuals -estimates population size -
Dispersion distribution of individuals in space, determines patterns of interaction among individuals
3 dispersion patterns -Clumped: presence of one indicates more there (pods of whales) -Regular: presence of one reduces probability of another nearby (territorial birds) -Random: equal chance of individual being anywhere (dandelions in a field)
Why so many humans? domestication of plants and animals, increasing crop and livestock yields through ongoing technological advances, plus medical advances
Equation for population N1 = N0 + (B – D) + (I – E)
Cohort life table tracks demographic events of a group of individuals born at the same time
Mortality proportion of each age class that die before reaching the next age class
Fecundity number of female offspring produced by each female, allows estimate of population’s potential for growth
3 patterns of survivorship curves Physiological, Ecological, and Maturational
Physiological (Type I) -high survivorship through adulthood -i.e. humans/large mammals -parental care & low fecundity
Ecological (Type II) Constant risk of mortality at all ages (i.e. most birds)
Maturational (type III) -low juvenile survivorship -many offspring and little parental care -many insects
life history strategy how an animal allocates time and energy among the various activities throughout its life
Semelparous animals that reproduce once, then die (i.e. salmon)
Iteroparous -animals that reproduce many times throughout their lives -typical of those that have high survivorship once they reach maturity
Guppies in Trinidad THM Predation favored early and frequent reproduction ->changing the guppy genotype.
per capita growth rate (r) r=b-d If birth rate exceeds death rate, r > 0, the population is growing
biotic potential rN, which represents the maximum growth rate a species may reach
intrinsic rate of increase -maximum value for r -rate of increase possible under ideal conditions -
carrying capacity (K) -number of individuals that can be supported in an environment indefinitely -determined by the availability of resources -logistic growth pattern
density-dependent factors -increase in proportion to population density -food supplies become limited -predators attracted to high densities of prey -pathogens spread more quickly
density-independent factors impact does not depend on population density, such as intense storms or cold periods
r-strategists life history strategies that allow for high intrinsic rate of increase
K-strategists life history strategies allow them to persist at or near the carrying capacity
3 factors that influence population density resource abundance (herbivores>), size of individuals (smaller>), time species has lived in an area (new area>)
metapopulation The larger population to which the subpopulations belong
Corridors -connections between patches in populations facilitate dispersal to maintain subpopulations -When habitat patches were connected by corridors, more species were able to maintain populations
Biological control use of natural predators to control pests
Chapter 57 <(o.o<) (>o.o)> <(0.0)>
Ecological community all species living/interacting in an area
Characterization of communities 1. Species composition—the number and kinds of species they contain. 2. The relative abundances of those species.
Photosynthetic Primary Production (PP) fixation of solar energy by autotrophs in an ecosystem
NPP = GPP – R GPP = total amount of energy fixed by all autotrophs in ecosystem NPP = energy left over after the autotrophs have met their energetic needs, i.e. respiration (R)
Trophic levels Primary producers -> Herbivores (Primary Consumers) -> Carnivores (secondary consumers) ->Detritovores (decomposers)
Omnivores Organisms that get food from more than one trophic level
Food webs interconnected food chains that show trophic interactions in a community.
Ecological Efficiency -the overall transfer of energy from one trophic level to the next -only approx. 10% of energy from one trophic level is transferred to the next
Limits of trophic levels -progressive energy loss limits trophic levels (mostly between 3-5) -each level has fewer species, less offspring, and bigger mass -
Trophic Cascade progression of indirect effects across successive trophic levels (i.e. the explosion of elk when wolves removed from Yellowstone) -other examples: bunny boom in Tasmania & zooplanktivores in ocean)
Ecosystem engineers organisms that influence ecosystem structure (i.e. beavers)
Keystone species A species that exerts influence out of proportion with its abundance (i.e. sea stars, sea otters, and freshwater bass)
Alpha diversity -within a single community or habitat; measured by counting species -same as species richness
Beta diversity between-site diversity; change in species composition from one habitat (or site) to another
Gamma diversity regional diversity over a range of communities in a larger geographic region
Field Example in Diversity Rivers: high species richness (alpha diversity) but low beta diversity (all rivers had same species) Ponds: high beta diversity Ditches: lowest alpha diversity, but many species were found in only one ditch- very high beta diversity
More diversity towards equator organisms had more time to diversify, more habitat types, more competition, and increased predation led to survival of rare species
species-area relationship mathematical relationship between the size of an area of habitat and the number of species that area contains
Immigration/extinction of island species influenced by... Distance from species pool and size of island
a disturbance an event that changes the survival rate of one or more species (i.e. windstorm that knocks down trees)
Succession Patterns of change in community composition following disturbance is called
3 types of succession Primary succession begins on sites that lack living organisms Secondary succession begins on sites where some organisms have survived Directional succession is characterized by a predictable progression of community assemblages
Species richness Environments that are rich in species are more efficient and prepared for environmental change
Monocultures -single species -how crops are grown -vulnerable to outbreaks of pests -polycultures (many crops) can reduce pests through more diversity
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